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Tree of Whispers, an end-game system that will provide you with frequently cycling world objectives and bounties that reward you with Legendary gear, experience, crafting materials, and more.
Blizzard desribes the Tree of Whispers as follows:
"...frequently cycling world objectives and bounties that reward you with Legendary gear, experience, crafting materials, Grim Favors and more".
Players will see "Whispers" throughout their travels around Sanctuary and completing these brings rewards of experience, gold, and Grim Favors.
Once 10 Grim Favours have been collected. they can be exchanged at the Tree of Whispers for a "bevy of loot" according to Blizzard.
The Whispers players see on their map will rotate frequently, with new ones becoming available throughout the day for them to track. Each Whisper will offer both a different set of rewards and experience for completing it.
Note, the Tree of Whispers content will not be available until a "specific chapter" of the main questline has been completed.