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What are Glyphs?
Glyphs are not to be confused with runes. Glyphs are specific to Diablo 4 and are items that can be found throughout Sanctuary. They are used specificaly in the Paragon Board.
On the Paragon boars there are special tiles in which a Glyph can be embed. When slotted, these Glyphs provide a bonus based on the number of active tiles within their radius.
Leveling up Glyphs
Glyphs can be levelled up by entering "some particulalry dangerous dungeons". Leveling up a Glyph extends the radius of their effect, allowing each one to draw power from or impart power to even more active tiles.